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Washington State University Molecular Plant Sciences

Phone 509-335-6198
FAX 509-335-8674
Assistant Professor, Crop and Soil Sciences


Ph.D., Washington State University, Crop Science, 2009
M.S., University of Idaho, Plant Science, 2006
B.S., University of Idaho, Plant Science, 2003


I lead the winter wheat breeding and genetics program at Washington State University.  My research is directed toward breeding improved wheat varieties for cropping systems in Washington state that incorporate diverse rotations and environments.  Our goal is to release high-yielding, disease resistant varieties with good end-use quality  that will maintain profitability and reduce the risk to growers.  New varieties are developed using a combination of traditional plant breeding methods, molecular marker technology, and biotechnology. Research efforts include identifying genetic solutions to winter wheat production problems, thereby enhancing the sustainability of the Washington wheat industry. Farmer participation is utilized in the development of new varieties and in research planning.  Current research topics involve developing water-use efficient wheat, adaptation of sensor based technology into wheat breeding programs, genetic understanding of end-use quality in wheat, and identification of new genes for stripe rust resistance.


Case, A., Y. Naruoka, X. Chen, K. Garland-Campbell, B. Zemetra, and A.H. Carter. (accepted) Mapping stripe rust resistance genes in a BrundageXCoda recombinant inbred line population. PlosONE

Higginbotham, R., P. Froese, and A.H. Carter. (in press) Tolerance of wheat (Triticum aestivum L.) seedlings to wireworm (Coleoptera: Elateridae). Journal of Economic Entomology.

Case, A., D. Skinner, K. Garland-Campbell, and A.H. Carter. (2014) Freezing tolerance-associated QTL in the Brundage x Coda wheat recombinant inbred line population.  Crop Science (First Look).

Carter, A.H., S.E. Cambron, H.W. Ohm, N. Bosque-Pèrez, and K.K. Kidwell. (2014) Identifying molecular markers associated with Hessian fly (Mayetiola destructor [Say]) resistance in the spring wheat (Tricitum aestivum) cultivar ‘Louise’. Crop Science (First Look).

Carter, A.H., S.S. Jones, G.B. Shelton, R. Higginbotham, S. Lyon, K. Balow, S. Guy, B. Baik, D.A. Engle, C. Morris, and X.M. Chen. 2012. Registration of ‘Otto’ wheat.  Journal of Plant Registrations 7:195-200.

Higginbotham, R., S.S. Jones, and A.H. Carter. 2013. Wheat cultivar performance and stability between no-till and conventional tillage systems in the Pacific Northwest of the United States. Sustainability 5:882-895.

Cavanagh, C.R., S. Chao, S. Wang, B.E. Huang, S. Stephen, S. Kiani, K. Forrest, C. Saintenac, G.L. Brown-Guedira, A. Akhunova, D. See, G. Bai, M. Pumphrey, L. Tomar, D. Wong, S. Kong, M. Reynolds, M. Lopez da Silva, H. Bockelman, L. Talbert, J.A. Anderson, S.Dreisigacker, P.S. Baenziger, A. Carter, V. Korzum, P.L. Morrell, J. Dubcovsky, M.K. Morell, M.E. Sorrells, M. Hayden, and E. Akhunov* (2013) Genome-wide comparative diversity uncovers multiple targets of selection for improvement in hexaploid wheat landraces and cultivars.  Proc Natl Acad Sci Online April 29, 2013 doi:10.1073/pnas.1217133110

Flowers, M., P.B. Hamm, A.H. Carter, and T.D. Murray* (2012) Reaction of winter wheat cultivars and breeding lines to soilborne wheat mosaic, 2011. Plant Disease Management Reports 6:CF025.

Lanning, S.P., P. Hucl, M. Pumphrey, A.H. Carter, P.F. Lamb, G.R. Carlson, D.M. Wichman, K.D. Kephart, D. Spaner, J.M. Martin, and L.E. Talbert* (2012) Agronomic performance of spring wheat as related to planting date and photoperiod response. Crop Science 52:1633-1639.

Poole, G.J., R.W. Smiley, T.C. Paulitz, A.H. Carter, D.R. See, and K. Garland-Campbell*  (2012) Identification of microsatellite markers Xgwm247 and Xgwm299 linked to quantitative trait loci for resistance to Fusarium crown rot (Fusarium pseudograminearum) in two spring wheat populations.  Theoretical and Applied Genetics 125:91-107.

Beecher, B.S.*, A.H. Carter, and D.R. See. 2012. Genetic mapping of a new family of seed-expressed polyphenol oxidase genes in wheat (Triticum aestivum L.).  Theoretical and Applied Genetics 124:1463-1473.

Carter, A.H.*, K. Garland-Campbell, C. Morris, and K.K. Kidwell. 2012. Chromosomes 3B and 4D are associated with several milling and baking quality traits in a soft white spring wheat (Triticum aestivum L.) population. Theoretical and Applied Genetics 124:1079-1096.

Carter, A.H.*, D.K. Santra, and K.K. Kidwell. 2012.  Assessment of the effect of the GPC-B1allele on senescence rate, grain protein concentration and mineral content in hard red spring wheat (Triticum aestivum L.) from the Pacific Northwest region of the USA. Plant Breeding 131:62-68.

Higginbotham, R.W, S.S. Jones, and A.H. Carter*.  2011. Adaptability of wheat cultivars to a late-planted no-till fallow production system. Sustainability 3:1224-1233.

Carter, A.H.*, K. Garland-Campbell, and K.K. Kidwell. 2011. Genetic mapping of quantitative trait loci associated with important agronomic traits in the spring wheat (Triticum aestivum L.) cross ‘Louise’ by ‘Penawawa’. Crop Science 51:84-95.